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In Large Carnivores and the Conservation of Biodiversity. No temptations, no regrets while gritting my teeth as we passed by the doughnut shop. This is in contrast to young foxes and coyotes, which may begin fighting even before the onset of play behavior. From there, the compound hormone-receptor diffuses into the nucleus, where it either alters the expression of genes [] or activates processes that send signals to other parts of the cell. National coat of arms Arms of dominion Ecclesiastical heraldry Burgher arms Civic heraldry Canting arms Attributed arms.

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There is also the risk that an intimate partner or child may come in contact with the application site and inadvertently dose himself or herself; children and women are highly sensitive to testosterone and can suffer unintended masculinization and health effects, even from small doses.

Injection is the most common method used by individuals administering AAS for non-medical purposes. The traditional routes of administration do not have differential effects on the efficacy of the drug. Studies indicate that the anabolic properties of AAS are relatively similar despite the differences in pharmacokinetic principles such as first-pass metabolism. However, the orally available forms of AAS may cause liver damage in high doses.

Known possible side effects of AAS include: Depending on the length of drug abuse, there is a chance that the immune system can be damaged. Most of these side-effects are dose-dependent, the most common being elevated blood pressure , especially in those with pre-existing hypertension. AAS have been shown to alter fasting blood sugar and glucose tolerance tests.

A number of severe side effects can occur if adolescents use AAS. For example, AAS may prematurely stop the lengthening of bones premature epiphyseal fusion through increased levels of estrogen metabolites , resulting in stunted growth. Other effects include, but are not limited to, accelerated bone maturation , increased frequency and duration of erections, and premature sexual development. AAS use in adolescence is also correlated with poorer attitudes related to health. Probably carcinogenic to humans.

Other side-effects can include alterations in the structure of the heart , such as enlargement and thickening of the left ventricle , which impairs its contraction and relaxation , and therefore reducing ejected blood volume. AAS use can cause harmful changes in cholesterol levels: AAS use in adolescents quickens bone maturation and may reduce adult height in high doses.

There are also sex-specific side effects of AAS. Development of breast tissue in males, a condition called gynecomastia which is usually caused by high levels of circulating estradiol , may arise because of increased conversion of testosterone to estradiol by the enzyme aromatase. This side-effect is temporary; the size of the testicles usually returns to normal within a few weeks of discontinuing AAS use as normal production of sperm resumes.

Female-specific side effects include increases in body hair , permanent deepening of the voice, enlarged clitoris , and temporary decreases in menstrual cycles. Alteration of fertility and ovarian cysts can also occur in females. Kidney tests revealed that nine of the ten steroid users developed a condition called focal segmental glomerulosclerosis , a type of scarring within the kidneys.

The kidney damage in the bodybuilders has similarities to that seen in morbidly obese patients, but appears to be even more severe. High doses of oral AAS compounds can cause liver damage. A review in CNS Drugs determined that "significant psychiatric symptoms including aggression and violence, mania , and less frequently psychosis and suicide have been associated with steroid abuse. Long-term steroid abusers may develop symptoms of dependence and withdrawal on discontinuation of AAS".

Recreational AAS use appears to be associated with a range of potentially prolonged psychiatric effects, including dependence syndromes, mood disorders , and progression to other forms of substance abuse, but the prevalence and severity of these various effects remains poorly understood.

Large-scale long-term studies of psychiatric effects on AAS users are not currently available. DSM-IV lists General diagnostic criteria for a personality disorder guideline that "The pattern must not be better accounted for as a manifestation of another mental disorder, or to the direct physiological effects of a substance e.

As a result, AAS users may get misdiagnosed by a psychiatrist not told about their habit. Affective disorders have long been recognised as a complication of AAS use. From the mids onward, the media reported "roid rage" as a side effect of AAS. A review determined that some, but not all, randomized controlled studies have found that AAS use correlates with hypomania and increased aggressiveness, but pointed out that attempts to determine whether AAS use triggers violent behavior have failed, primarily because of high rates of non-participation.

Compared with individuals that did not use steroids, young adult males that used AAS reported greater involvement in violent behaviors even after controlling for the effects of key demographic variables, previous violent behavior, and polydrug use. The drug response was highly variable. The mechanism of these variable reactions could not be explained by demographic, psychological, laboratory, or physiological measures.

A study of two pairs of identical twins, in which one twin used AAS and the other did not, found that in both cases the steroid-using twin exhibited high levels of aggressiveness, hostility, anxiety, and paranoid ideation not found in the "control" twin. The relationship between AAS use and depression is inconclusive. There have been anecdotal reports of depression and suicide in teenage steroid users, [] but little systematic evidence.

A review found that AAS may both relieve and cause depression, and that cessation or diminished use of AAS may also result in depression, but called for additional studies due to disparate data.

Androgens such as testosterone , androstenedione and dihydrotestosterone are required for the development of organs in the male reproductive system , including the seminal vesicles , epididymis , vas deferens , penis and prostate. The pharmacodynamics of AAS are unlike peptide hormones. However, as fat-soluble hormones, AAS are membrane-permeable and influence the nucleus of cells by direct action. The pharmacodynamic action of AAS begin when the exogenous hormone penetrates the membrane of the target cell and binds to an androgen receptor AR located in the cytoplasm of that cell.

From there, the compound hormone-receptor diffuses into the nucleus, where it either alters the expression of genes [] or activates processes that send signals to other parts of the cell. The effect of AAS on muscle mass is caused in at least two ways: It has been hypothesized that this reduction in muscle breakdown may occur through AAS inhibiting the action of other steroid hormones called glucocorticoids that promote the breakdown of muscles. As their name suggests, AAS have two different, but overlapping, types of effects: Some examples of the anabolic effects of these hormones are increased protein synthesis from amino acids , increased appetite, increased bone remodeling and growth, and stimulation of bone marrow , which increases the production of red blood cells.

Through a number of mechanisms AAS stimulate the formation of muscle cells and hence cause an increase in the size of skeletal muscles , leading to increased strength.

The androgenic effects of AAS are numerous. Depending on the length of use, the side effects of the steroid can be irreversible. Processes affected include pubertal growth, sebaceous gland oil production, and sexuality especially in fetal development. Some examples of virilizing effects are growth of the clitoris in females and the penis in male children the adult penis size does not change due to steroids [ medical citation needed ] , increased vocal cord size, increased libido , suppression of natural sex hormones , and impaired production of sperm.

Men may develop an enlargement of breast tissue, known as gynecomastia, testicular atrophy, and a reduced sperm count. Compounds with a high ratio of androgenic to an anabolic effects are the drug of choice in androgen-replacement therapy e. This disassociation is less marked in humans, where all AAS have significant androgenic effects. A commonly used protocol for determining the androgenic: The VP weight is an indicator of the androgenic effect, while the LA weight is an indicator of the anabolic effect.

Two or more batches of rats are castrated and given no treatment and respectively some AAS of interest. Animal studies also found that fat mass was reduced, but most studies in humans failed to elucidate significant fat mass decrements.

The effects on lean body mass have been shown to be dose-dependent. Both muscle hypertrophy and the formation of new muscle fibers have been observed. The hydration of lean mass remains unaffected by AAS use, although small increments of blood volume cannot be ruled out. The upper region of the body thorax, neck, shoulders, and upper arm seems to be more susceptible for AAS than other body regions because of predominance of ARs in the upper body.

After drug withdrawal, the effects fade away slowly, but may persist for more than 6—12 weeks after cessation of AAS use. Overall, the exercise where the most significant improvements were observed is the bench press. The measurement of the dissociation between anabolic and androgenic effects among AAS is based largely on a simple although arguably unsophisticated and outdated model involving rat tissue bioassays. The intracellular metabolism theory explains how and why remarkable dissociation between anabolic and androgenic effects can occur despite the fact that these effects are mediated through the same signaling receptor, and of course why dissociation is invariably incomplete.

An animal study found that two different kinds of androgen response elements could differentially respond to testosterone and DHT upon activation of the AR. Changes in endogenous testosterone levels may also contribute to differences in myotrophic—androgenic ratio between testosterone and synthetic AAS.

Testosterone can be metabolized by aromatase into estradiol , and many other AAS can be metabolized into their corresponding estrogenic metabolites as well. The major effect of estrogenicity is gynecomastia woman-like breasts. AAS are androstane or estrane steroids. As well as others such as 1-dehydrogenation e. The most commonly employed human physiological specimen for detecting AAS usage is urine, although both blood and hair have been investigated for this purpose.

The AAS, whether of endogenous or exogenous origin, are subject to extensive hepatic biotransformation by a variety of enzymatic pathways. The primary urinary metabolites may be detectable for up to 30 days after the last use, depending on the specific agent, dose and route of administration.

A number of the drugs have common metabolic pathways, and their excretion profiles may overlap those of the endogenous steroids, making interpretation of testing results a very significant challenge to the analytical chemist. Methods for detection of the substances or their excretion products in urine specimens usually involve gas chromatography—mass spectrometry or liquid chromatography-mass spectrometry.

The use of gonadal steroids pre-dates their identification and isolation. Medical use of testicle extract began in the late 19th century while its effects on strength were still being studied. In the s, it was already known that the testes contain a more powerful androgen than androstenone , and three groups of scientists, funded by competing pharmaceutical companies in the Netherlands, Germany, and Switzerland, raced to isolate it.

The chemical synthesis of testosterone was achieved in August that year, when Butenandt and G. Wettstein, announced a patent application in a paper "On the Artificial Preparation of the Testicular Hormone Testosterone Androstenoneol.

Clinical trials on humans, involving either oral doses of methyltestosterone or injections of testosterone propionate , began as early as Kennedy was administered steroids both before and during his presidency. The development of muscle-building properties of testosterone was pursued in the s, in the Soviet Union and in Eastern Bloc countries such as East Germany, where steroid programs were used to enhance the performance of Olympic and other amateur weight lifters.

In response to the success of Russian weightlifters, the U. The new steroid was approved for use in the U. It was most commonly administered to burn victims and the elderly. The drug's off-label users were mostly bodybuilders and weight lifters. Although Ziegler prescribed only small doses to athletes, he soon discovered that those having abused Dianabol suffered from enlarged prostates and atrophied testes. Three major ideas governed modifications of testosterone into a multitude of AAS: Androgens were discovered in the s and were characterized as having effects described as androgenic i.

Although anabolic steroid was originally intended to specifically describe testosterone-derived steroids with a marked dissociation of anabolic and androgenic effect, it is applied today indiscriminately to all steroids with AR agonism-based anabolic effects regardless of their androgenic potency, including even non-synthetic steroids like testosterone itself.

Infections by the tapeworm Echinococcus granulosus in ungulate populations tend to increase in areas with high wolf densities, as wolves can shed Echinoccocus eggs in their feces onto grazing areas. Wolves can carry over 30 roundworm species, though most roundworm infections appear benign, depending on the number of worms and the age of the host.

Ancylostoma caninum attaches itself on the intestinal wall to feed on the host's blood, and can cause hyperchromic anemia , emaciation, diarrhea , and possibly death. Toxocara canis , a hookworm known to infect wolf pups in utero, can cause intestinal irritation, bloating, vomiting, and diarrhea.

Wolves can tolerate low levels of Dirofilaria immitis for many years without showing any ill effects, though high levels can kill wolves through cardiac enlargement and congestive hepatopathy. Wolves probably become infected with Trichinella spiralis by eating infected ungulates. Nicolla skrjabini , Macrocantorhynchus catulinus , and Moniliformis moniliformis.

Human presence appears to stress wolves, as seen by increased cortisol levels in instances such as snowmobiling near their territory.

Old English literature contains several instances of Anglo-Saxon kings and warriors taking on wulf as a prefix or suffix in their names. Wolf-related names were also common among pre-Christian Germanic warriors: Ancient Greek literature is similar: Autolycus "the wolf itself" , Lycurgus "wolf-work". The Latin for "female prostitute" is lupa , and the most famous brothel in Pompeii was the Lupanar. The wolf is a common motif in the foundational mythologies and cosmologies of peoples throughout Eurasia and North America corresponding to the historical extent of the habitat of the gray wolf.

The obvious attribute of the wolf is its nature of a predator , and correspondingly it is strongly associated with danger and destruction, making it the symbol of the warrior on one hand, and that of the devil on the other.

The modern trope of the Big Bad Wolf is a development of this. The wolf holds great importance in the cultures and religions of the nomadic peoples, both of the Eurasian steppe and of the North American Plains. In many cultures, the identification of the warrior with the wolf totemism gave rise to the notion of Lycanthropy , the mythical or ritual identification of man and wolf.

Aesop featured wolves in several of his fables , playing on the concerns of Ancient Greece 's settled, sheep-herding world. His most famous is the fable of The Boy Who Cried Wolf , which is directed at those who knowingly raise false alarms, and from which the idiomatic phrase " to cry wolf " is derived.

Some of his other fables concentrate on maintaining the trust between shepherds and guard dogs in their vigilance against wolves, as well as anxieties over the close relationship between wolves and dogs. Although Aesop used wolves to warn, criticize and moralize about human behavior, his portrayals added to the wolf's image as a deceitful and dangerous animal.

Although portrayed as loyal, honest and moral, Isengrim is forever the victim of Reynard's wit and cruelty, often dying at the end of each story. The tale of Little Red Riding Hood , first written in by Charles Perrault , is largely considered to have had more influence than any other source of literature in forging the wolf's negative reputation in the western world.

The wolf in this story is portrayed as a potential rapist , capable of imitating human speech. Tolstoy's War and Peace and Chekhov's Peasants both feature scenes in which wolves are hunted with hounds and borzois.

Although credited with having changed popular perceptions on wolves by portraying them as loving, cooperative and noble, it has been criticized for its idealization of wolves and its factual inaccuracies. The wolf is a frequent charge in English armory. It is illustrated as a supporter on the shields of Lord Welby , Rendel , and Viscount Wolseley , and can be found on the coat of arms of Lovett and the vast majority of the Wilsons and Lows.

The demi-wolf is a common crest , appearing in the arms and crests of members of many families, including that of the Wolfes , whose crest depicts a demi-wolf holding a crown in its paws, in reference to the assistance the family gave to Charles II during the battle of Worcester. Wolf heads are common in Scottish heraldry , particularly in the coats of Clan Robertson and Skene. The wolf is the most common animal in Spanish heraldry , and is often depicted as carrying a lamb in its mouth, or across its back.

It is the unofficial symbol of the spetsnaz , and serves as the logo of the Turkish Grey Wolves. Livestock depredation has been one of the primary reasons for hunting wolves, and can pose a severe problem for wolf conservation: Being the most abundant carnivores, feral and free-ranging dogs have the greatest potential to compete with wolves. A review of the studies in the competitive effects of dogs on sympatric carnivores did not mention any research on competition between dogs and wolves.

Wolves kill dogs on occasion, with some wolf populations relying on dogs as an important food source. Wolves may display unusually bold behavior when attacking dogs accompanied by people, sometimes ignoring nearby humans. Large hunting dogs such as Swedish elkhounds are more likely to survive wolf attacks because of their better ability to defend themselves. Although the numbers of dogs killed each year are relatively low, it induces a fear of wolves entering villages and farmyards to take dogs.

In many cultures, there are strong social and emotional bonds between humans and their dogs that can be seen as family members or working team members. The loss of a dog can lead to strong emotional responses with demands for more liberal wolf hunting regulations. Dogs that are employed to guard sheep help to mitigate human—wolf conflicts, and are often proposed as one of the non-lethal tools in the conservation of wolves.

The fear of wolves has been pervasive in many societies, though humans are not part of the wolf's natural prey. Predatory attacks attacks by wolves treating humans as food may be preceded by a long period of habituation , in which wolves gradually lose their fear of humans. The victims are repeatedly bitten on the head and face, and are then dragged off and consumed, unless the wolves are driven off.

Such attacks typically occur only locally, and do not stop until the wolves involved are eliminated. Predatory attacks can occur at any time of the year, with a peak in the June—August period, when the chances of people entering forested areas for livestock grazing or berry and mushroom picking increase, [24] though cases of non-rabid wolf attacks in winter have been recorded in Belarus , Kirov and Irkutsk oblasts, Karelia and Ukraine.

The majority of victims of predatory wolf attacks are children under the age of 18 and, in the rare cases where adults are killed, the victims are almost always women. Cases of rabid wolves are low when compared to other species, as wolves do not serve as primary reservoirs of the disease, but can be infected by animals such as dogs, jackals and foxes.

Incidents of rabies in wolves are very rare in North America, though numerous in the eastern Mediterranean , Middle East and Central Asia. Wolves apparently develop the "furious" phase of rabies to a very high degree which, coupled with their size and strength, makes rabid wolves perhaps the most dangerous of rabid animals, [24] with bites from rabid wolves being 15 times more dangerous than those of rabid dogs.

Most rabid wolf attacks occur in the spring and autumn periods. Unlike with predatory attacks, the victims of rabid wolves are not eaten, and the attacks generally only occur on a single day. The victims are chosen at random, though the majority of cases involve adult men. In the half-century up to , there were eight fatal attacks in Europe and Russia, and more than in south Asia. Wolves are difficult to hunt because of their elusiveness, sharp senses, high endurance, and ability to quickly incapacitate and kill hunting dogs.

This method relies heavily on the wolf's fear of human scents, though it can lose its effectiveness when wolves become accustomed to the smell. Wild wolves are sometimes kept as exotic pets and, in some rarer occasions, as working animals. Although closely related to domesticated dogs , wolves do not show the same tractability as dogs in living alongside humans, and generally, much more work is required in order to obtain the same amount of reliability.

From Wikipedia, the free encyclopedia. Redirected from Gray wolf. This article is about the gray wolf, canis lupus , and its subspecies. For other species of wolf and other uses, see Wolf disambiguation. For other uses, see Grey Wolf disambiguation. Middle Pleistocene — present , years BP [1]. Linnaeus , [3]. Subspecies of Canis lupus. Evolution of the wolf. List of gray wolf populations by country.

Wolves in folklore, religion and mythology. Wolf attacks on humans and List of wolf attacks. Wolf hunting and Wolf hunting with dogs. Human uses of hunted wolves. Wolves as pets and working animals. However, neither dogs nor dingoes are referred to as gray wolves. Throughout this article, the term "wolf" will be used to refer collectively to naturally occurring subspecies, especially the nominate subspecies, Canis lupus lupus.

This terminology was first used in by Rudolf Schenkel of the University of Basel , who based his findings on researching the behavior of captive gray wolves. This view on gray wolf pack dynamics was later popularized by L.

David Mech in his book The Wolf. He formally disavowed this terminology in , explaining that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves.

Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and its offspring of the previous 1—3 years.

Bulletin of the American Museum of Natural History. International Union for Conservation of Nature. Tomus I in Latin 10 ed. Retrieved November 23, Gray wolf Canis lupus and allies", in Feldhamer, George A. Wild Mammals of North America: Grey wolf Canis lupus Linnaeus, , pp. Foxes, Wolverhampton Wanderers F.

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Its Domestication and Behavior. The cautionary case of Bulgarian wolves". The Fear of Wolves: Norsk institutt for naturforskning NINA. A contribution to the study of wolf-dog hybridization patterns". Mammalian Biology - Zeitschrift für Säugetierkunde. Archived from the original PDF on September 24, Retrieved June 2, A new profile for the wolf. Journal of Vertebrate Paleontology.

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African palm civet N. Angolan slender mongoose G. Ethiopian dwarf mongoose H. Family Viverridae includes Civets. It's my review of Nutrisystem, and I believe you will find it one of the most interesting Nutrisystem reviews you will ever read. Does Nutrisystem work for everyone? We are all different and not everyone will have the same results.

I hope that my story might help someone else suffering from weight issues. Growing up, I never had a weight problem. In fact I was always underweight. When I married at age 22, I was only lbs. I wore a size zero!

It seemed like every month my weight increased. When my daughter left for college, worry and stress led to my binge eating. I love to eat. Bread is my weakness. In result, every time I would go shopping for new clothes, I found myself reaching for a larger size.

My weight gain eventually started causing health problems. Eventually my knees started giving me problems. They hurt a lot therefore making it uncomfortable for me to enjoy an afternoon walk. I also started suffering from acid reflux. Sleeping at night was uncomfortable due to the discomfort and chest pains. I even noticed poor circulation in my arms and legs. My self-esteem became low and had a bad attitude that I would never be small again. I was miserable most of the time, except when I was eating of course.

I knew I needed to go on a diet. As diabetes runs in my family, I knew I was setting myself up for that path. As a matter of fact, I would start some kind of diet every Monday. I tried several different diets on my own. I cut out carbs, sweets, and calorie intakes. Other diets were boring and felt like a waste of time since I never felt any changes.

I would often kid around with my husband by telling him that he was actually only married to half of me. I finally decided to take control and do something about my weight. It was January 5, I mentioned to my husband that I was going to try Nutrisystem.

I was spectacle since no other diets seem to work. I headed to my local Walmart and purchased a Nutrisystem five day weight loss kit which included fifteen entrees and five desserts. I was a bit embarrassed as I walked through the store carrying a box of Nutrisystem. I felt like people were staring as I left the store.

He believed me this time and even made a choice to diet with me. He wanted to be supportive. One way was by not having tempting foods in the house. This was a big plus. No temptations, no regrets while gritting my teeth as we passed by the doughnut shop. When I got home and opened my Nutrisystem box, there were a few guidelines to follow to achieve my goal. Wait, do I have a goal? I never really thought about setting a goal, I just wanted to lose weight. The first morning I grabbed my cup of coffee and sat down to a tiny little Nutrisystem muffin and a glass of fusion energy juice.

I was hungry all day. Nutrisystem labels snacks as either Power Fuels or Smart Carbs. The kit included a list of this type of foods and servings allowed for each. The truth is, I was eating the wrong things to achieve any positive results. I added healthy snacks between my meals such as fresh avocado, yogurt and mixed nuts. The days got easier. By the third day of sticking to the plan which was a big accomplishment for me , I was already feeling more energy and less bloating.

This motivated me to do another week. Weeks went on and I continued following the Nutrisystem diet. So, how long does Nutrisystem take to work? It works within the first few days. Each day I made sure to drink at least 64 oz of water and light exercise, which usually included twenty minutes of hula hooping. With the help of my Fitbit, I made a goal to get in at least steps in per day. I used the MyfitnessPal app to keep a log of my daily food and calorie intake.

Here I was able to make other friends from a Nutrisystem support group who were reaching for the same goals. The weekends were challenging as we chose to go out to our favorite restaurants. I ordered off the Lite menu or A La Carte. The hardest part was avoiding the chips and dip at the Mexican restaurant.

This usually resulted in me actually sitting on my hands. Yes the struggle was real here. The second week was easier. I finally set a goal to lose 30 lbs. I really wanted to get into a new Easter outfit. I was still buying Nutrisystem by the week from Walmart. Does Nutrisystem from Walmart or Costco work? It sure did for me. I looked forward to my next meal. I got hungry between meals sometimes.

This is when I would snack on foods such as almonds, kale chips or nonfat yogurts. I purchased some Nutrisystem shakes to use between meals also. They are delicious and will knock out the hungry feelings or cravings. I only lost 2 lbs the second week but it was still a loss. By the third week, I noticed my health was improving.

I no longer suffered from heartburn and acid reflux. I felt good and best of all I was losing weight. By now, I knew this was the diet for me and was ready to keep going.

I decided to give in and purchase through Nutrisystem. While I was online ordering I chatted with a representative about my plan options and payment concerns. She was very informative as she helped me make my first purchase. Once I became a member, there were tools on the website that helped me achieve my goal. I entered my height and weight, how many pounds I wanted to lose and it set a calorie goal for me. I needed calories a day to achieve my desired weight loss goal of 30 lbs.

I was able to choose days worth of food from the menu. A couple days later a large box showed up at my door. I stocked my pantry and the journey continued. After one month the whole diet thing was easier as I was no longer craving foods like before. The cravings were not completely gone but walking away from a piece of cake or pizza was no longer a challenge. I had my own Nutrisystem goodies such as chocolate, pizza, potatoes, muffins and more.

I stepped on the scales and smiled as I had lost a total of 11 lbs in one month. This earned me a Nutribear. Nutrisystem will award a new bear for every 10 lbs. New eating habits were not my only struggle. To accomplish success, this required me to give up food and wine outings with friends.